Everything about The Jaw totally explained
The
jaw is either of the two opposable structures forming, or near the entrance to, the
mouth.
The term
jaws is also broadly applied to the whole of the structures constituting the vault of the mouth and serving to open and close it and is part of the
body plan of most animals.
Arthropods
In
arthropods, the jaws are
chitinous and oppose laterally, and may consist of
mandibles,
chelicerae, or loosely,
pedipalps.
Their function is fundamentally for food acquisition, conveyance to the mouth, and/or initial processing (
mastication or
chewing).
Vertebrates
In most
vertebrates, the jaws are
bony or
cartilaginous and oppose vertically, comprising an
upper jaw and a
lower jaw.
Bones of the jaw
In vertebrates, the lower jaw, dentary or
mandible is the mobile component that articulates at its posterior processes, or
rami (singular
ramus), with the temporal bones of the
skull on either side; the word
jaw used in the singular typically refers to the lower jaw.
The upper jaw or
maxilla is more or less fixed with the skull and is composed of two bones, the
maxillae, fused intimately at the
median line by a
suture; incomplete closure of this suture and surrounding structures may be involved in the malformation known as
cleft palate.
The maxillary bones form parts of the roof of the mouth, the floor and sides of the
nasal cavity, and the floor of the
orbit or
eye socket.
The jaws typically accommodate the
teeth or form the bases for the attachment of a
beak.
The jaw in fish and amphibians
The vertebrate jaw probably originally evolved in the
Silurian period and appeared in the
Placoderm fish which further diversified in the
Devonian. Jaws are thought to derive from the pharyngeal arches that support the
gills in fish. The two most anterior of these arches are thought to have become the jaw itself and the hyoid arch, which braces the jaw against the braincase and increases mechanical efficiency. While there's no fossil evidence directly to support this theory, it makes sense in light of the numbers of pharyngeal arches that are visible in extant jawed (the
Gnathostomes), which have seven arches, and primitive jawless vertebrates (the
Agnatha), which have nine.
It is thought that the original selective advantage garnered by the jaw wasn't related to feeding, but to increased respiration efficiency. The jaws were used in the
buccal pump (observable in modern fish and
amphibians) that pumps water across the gills of fish or air into the lungs in the case of amphibians. Over evolutionary time the more familiar use of jaws (to humans), in feeding, was selected for and became a very important function in vertebrates.
The jaw in reptiles
In
reptiles, the
mandible is made up of five bones. In the evolution of
mammals, four of these bones were reduced in size and incorporated into the ear. In their reduced form, they're known as the
malleus and
incus; along with the more ancient
stapes, they're the
ossicles. This adaptation is advantageous, not only because a one-bone jaw is stronger, but also because the malleus and incus improve hearing. (However, reptiles tend to swallow prey whole because their pace of digestion is different than mammals, so multiple jaw bones may allow flexibility to expand the jaws around prey.)
Further Information
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